1. Department of Plant and Microbial Biology University of Minnesota‐Twin Cities 1445 Gortner Avenue St. Paul Minnesota 55108‐1095 USA
2. Systematics, Biodiversity and Evolution of Plants, Department of Biology I, Ludwig‐Maximilians‐Universität München Menzinger Strasse 67 80638 Munich Germany
3. College of Life Science Henan Agricultural University 63 Nongye Road Zhengzhou Henan 450002 China
4. Biological Sciences Department California Polytechnic State University, 1 Grand Avenue, San Luis Obispo California 93407 USA
5. Department of Biological Sciences Binghamton University (State University of New York), 4400 Vestal Parkway E, Binghamton New York 13902 USA
6. Department of Botany & Rocky Mountain Herbarium University of Wyoming, 1000 E. University Avenue, Laramie Wyoming 82071 USA