1. Biology Department, Colorado College, 14 E. Cache La Poudre, Colorado Springs, Colorado 80903 USA;
2. California Department of Food and Agriculture, 1220 N Street, Sacramento, California 95814 USA;
3. Department of Ecology, Evolution and Marine Biology, University of California, Santa Barbara, California 93106 USA; and
4. Department of Biology, Box 355325, University of Washington, Seattle, Washington 98195 USA