1. Marine Science Institute University of California Santa Barbara California 93106 USA
2. Institute of Ecology College of Urban and Environmental Sciences Key Laboratory for Earth Surface Processes of the Ministry of Education Peking University Beijing 100871 China
3. Bren School of Environmental Science University of California Santa Barbara California 93106 USA
4. Western Ecological Research Center U.S. Geological Survey at Marine Science Institute University of California Santa Barbara California 93106 USA