1. National Research Institute of Brewing; Higashihiroshima Japan
2. Graduate School of Biosphere Sciences; Hiroshima University; Higashihiroshima Japan
3. Hyogo Prefectural Technology Center for Agriculture, Forestry and Fishers; Kato Hyogo Japan
4. Department of Biological Resource Sciences; Akita Prefectural University; Nakano Shimoshinjyo Akita Japan