1. Department of OceanographyMarine Research Institute, University of Cape TownRondebosch South Africa
2. Southern Ocean Carbon and Climate ObservatoryCSIRStellenbosch South Africa
3. Now at Department of Marine SciencesUniversity of GothenburgGothenburg Sweden
4. Physical Oceanography DepartmentWoods Hole Oceanographic InstitutionWoods Hole Massachusetts USA