1. Department of Animal Ecology and Tropical Biology Biocenter University of Würzburg Würzburg Germany
2. Unit of Environmental Sciences and Management North West University Potchefstroom South Africa
3. Department of Plant Systematics University of Bayreuth Bayreuth Germany
4. Department Arthropoda Zoological Research Museum Alexander Koenig Bonn Germany
5. Bundesamt für Naturschutz Bonn Germany