1. Germplasm Bank of Wild Species Kunming Institute of Botany, Chinese Academy of Sciences Kunming Yunnan China
2. CAS Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany Chinese Academy of Sciences Kunming Yunnan China
3. Kunming College of Life Science University of Chinese Academy of Sciences Kunming Yunnan China
4. Department of Biology, College of Letters and Sciences Columbus State University, University System of Georgia Columbus Georgia
5. Key Laboratory of Economic Plants and Biotechnology Kunming Institute of Botany, Chinese Academy of Sciences Kunming Yunnan China
6. Royal Botanic Garden Edinburgh Edinburgh UK