1. Hubei Key Laboratory of Quality Control of Characteristic Fruits and Vegetables, College of Life Sciences and Technology, Hubei Engineering University, Xiaogan 432000, China
2. School of Ecology and Environment, Tibet University, Lhasa 850000, China
3. Agriculture and Animal Husbandry Comprehensive Service Center, Jiangrang Township, Cuoqin County, Ngari 859000, China
4. College of Life Science, Wuhan University, Wuhan 430072, China
5. Key Laboratory of Biodiversity and Environment on the Qinghai-Tibetan Plateau, Ministry of Education, Tibet University, Lhasa 850000, China