1. Department of Geography; University of Southampton; Highfield Southampton S017 1BJ UK
2. Soils, Agriculture and Water; ADAS; Woodthorne, Wergs Road Wolverhampton WV6 8TQ, West Midlands UK
3. Tyne Rivers Trust; Horesley Northumberland NE15 OPA UK
4. School of Biological and Chemical Sciences; Queen Mary University of London; Mile End Road London E1 4NS UK
5. Centre for Ecology and Hydrology; Crowmarsh Gifford Wallingford Oxon UK