1. Department of Biological Science, College of Science; Sungkyunkwan University; Suwon South Korea
2. Univ. Lille, CNRS, Univ. Littoral Cote d'Opale, UMR 8187, LOG; Laboratoire d'Océanologie et de Géosciences; Wimereux France
3. Department of Marine Bioscience, College of Life Sciences; Gangneung-Wonju National University; Gangneung South Korea