1. Department of Biological Sciences, College of Science Sungkyunkwan University Suwon South Korea
2. Division of Life Sciences Korea Polar Research Institute Incheon South Korea
3. Laboratoire d'Océanologie et de Géosciences, Station marine de Wimereux, CNRS, UMR 8187 LOG Université de Lille, Université du Littoral Côte d'Opale Lille France
4. Department of Marine Resource Development, College of Life Sciences Gangneung‐Wonju National University Gangneung South Korea
5. Graduate School of Fisheries and Environmental Sciences Nagasaki University Nagasaki Japan
6. Organization for Marine Science and Technology Nagasaki University Nagasaki Japan