1. Department of Fisheries and Wildlife College of Agriculture and Natural Resources Michigan State University East Lansing Michigan 48824 USA
2. Department of Fisheries and Wildlife College of Agriculture and Natural Resources and Department of Pathobiology and Diagnostic Investigation College of Veterinary Medicine Michigan State University East Lansing Michigan 48824 USA
3. U.S. Department of Agriculture Agricultural Research Service National Center for Cool and Cold Water Aquaculture Kearneysville West Virginia 25430 USA
4. Quantitative Fisheries Center Department of Fisheries and Wildlife College of Agriculture and Natural Resources Michigan State University East Lansing Michigan 48824 USA