1. Institute for Plant Nutrition and Soil Science, CAU Kiel; Hermann-Rodewald-Str. 2 24118 Kiel Germany
2. Institute of Soil Landscape Research, Leibnitz Centre for Agricultural Landscape Research; Eberswalder Str. 84 15374 Müncheberg Germany
3. School of Biological Sciences; University of Aberdeen; Aberdeen AB24 3UU UK