1. Molecular Plant Nutrition, University of Hohenheim, 70593 Stuttgart, Germany
2. Institute for Biology, Humboldt University Berlin, Chausseestrasse 117, 10115 Berlin, Germany
3. Scottish Crop Research Institute, Invergowrie, Dundee, DD2 5DA, Scotland UK
4. School of Biological and Environmental Sciences, University of Stirling, Stirling FK9 4LA, Scotland UK
5. Leibniz Institute for Plant Genetics and Crop Plant Research, Corrensstr. 3, 06466 Gatersleben, Germany