1. Food Research Institute, NARO, 2-1-12, Kannondai, Tsukuba, Ibaraki 305-8642, Japan
2. Institute of Fruit Tree and Tea Science, NARO, 2-1 Fujimoto, Tsukuba, Ibaraki 305-0852, Japan
3. Graduate School of Horticulture, Chiba University, 648 Matsudo, Chiba 271-8510, Japan