1. Chengdu Institute of Biology, Chinese Academy of Sciences, Chengdu 610041, China
2. Nature Conservation of Snake Island and Laotieshan Mountain, Dalian 116041, China
3. Department of Integrative Biology, University of Guelph, Guelph, Ontario N1G 2W1, Canada
4. Key Laboratory of Vertebrate Evolution and Human Origins of Chinese Academy of Sciences, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Science, Beijing 100044, China
5. University of Chinese Academy of Sciences, Beijing 100049, China
6. College of life sciences and food engineering, Yibin University, Yibin 644007, China