1. Department of Biological Sciences National University of Singapore Singapore Singapore
2. Institute for Biodiversity and Ecosystem Dynamics University of Amsterdam Amsterdam the Netherlands
3. State Key Laboratory of Tibetan Plateau Earth System, Resources and Environment (TPESRE) Institute of Tibetan Plateau Research Chinese Academy of Sciences Beijing China
4. School of Geographical Sciences University of Bristol Bristol United Kingdom
5. Royal Botanic Gardens, Kew Richmond United Kingdom
6. Palynova Limited Littleport United Kingdom
7. Department of Earth Sciences Royal Holloway University of London Egham United Kingdom