1. Key Laboratory of Mariculture, Ministry of Education, College of Fisheries Ocean University of China Qingdao China
2. Institute of Evolution and Marine Biodiversity Ocean University of China Qingdao China
3. Shanghai Universities Key Laboratory of Marine Animal Taxonomy and Evolution Shanghai Ocean University Shanghai China
4. Engineering Research Center of Environmental DNA and Ecological Water Health Assessment Shanghai Ocean University Shanghai China
5. Department of Life Sciences Natural History Museum London UK