1. Laboratory of Protozoology; Institute of Evolution & Marine Biodiversity; Ocean University of China; Qingdao 266003 China
2. College of Fisheries and Life Science; Shanghai Ocean University; Shanghai 201306 China
3. Department of Life Sciences; Natural History Museum; Cromwell Road London SW7 5BD UK
4. Zoology Department; College of Science; King Saud University; Riyadh 11451 Saudi Arabia
5. Laboratory for Marine Biology and Biotechnology; Qingdao National Laboratory for Marine Science and Technology; Qingdao 266003 China