1. College of Fisheries & Key Laboratory of Mariculture, Ministry of Education, Ocean University of China , Qingdao 266003 , China
2. Institute of Evolution and Marine Biodiversity, Ocean University of China , Qingdao 266003 , China
3. Zoology Department, College of Science, King Saud University , Riyadh 11451 , Saudi Arabia
4. Department of Life Sciences, Natural History Museum , London , UK