1. Institute of Crop Science, National Agriculture and Food Research Organization (NARO) 2‐1‐2 Kannondai, Tsukuba Ibaraki 305‐8602 Japan
2. Graduate School of Horticulture Chiba University 648 Matsudo, Matsudo Chiba 271‐8510 Japan
3. Key Laboratory of Aquatic Genomics, Ministry of Agriculture and Rural Affairs, and Beijing Key Laboratory of Fishery Biotechnology Chinese Academy of Fishery Sciences Beijing 100141 China
4. Crop Research Institute, Shandong Academy of Agricultural Sciences/National Engineering Research Center of Wheat and Maize/Shandong Technology Innovation Center of Wheat Jinan 252100 China