1. Asian Natural Environmental Science Center; The University of Tokyo; 1-1-1 Yayoi Bunkyo Tokyo 113-8657 Japan
2. Biological Institute; Section of Marine Biology; University of Copenhagen; Universitetsparken 4 DK-2100 Copenhagen K Denmark
3. Graduate School of Fisheries and Environmental Sciences; Nagasaki University; 1-14 Bunkyo Nagasaki 852-8521 Japan
4. Research Institute for Marine Fisheries; 222 Le Hai Hai Phong Vietnam