1. College of Fisheries, Guangdong Ocean University, Zhanjiang 524088, China
2. Guangdong Laboratory of Marine Ecology Environment Monitoring and Warning, Zhanjiang 524088, China
3. CAS Key Laboratory of Marine Ecology and Environmental Sciences, Institute of Oceanology, Chinese Academy of Sciences, Qingdao 266071, China
4. Functional Laboratory of Marine Ecology and Environmental Science, Laoshan Laboratory, Qingdao 266237, China
5. Third Institute of Oceanography, Ministry of Natural Resources, Xiamen 361005, China
6. CAS Key Laboratory of Experimental Marine Biology, Institute of Oceanology, Chinese Academy of Sciences, Qingdao 266071, China