1. CAS Key Laboratory of Marine Ecology and Environmental Sciences, Institute of Oceanology, Chinese Academy of Sciences, Qingdao 266071, China
2. Laboratory for Marine Ecology and Environmental Science, Laoshan Laboratory for Marine Science and Technology, Qingdao 266237, China
3. Center for Ocean Mega-Science, Chinese Academy of Sciences, Qingdao 266071, China
4. Current address: Department of Aquaculture, College of Fisheries, Guangdong Ocean University, Zhanjiang 524088, China
5. Third Institute of Oceanography, Ministry of Natural Resources, Xiamen 361005, China
6. University of Chinese Academy of Sciences, Beijing 100049, China