1. Institute of Earth Sciences University of Lausanne 1015 Lausanne Switzerland
2. Department of Earth Sciences, Palaeobiology Uppsala University Uppsala 752 36 Sweden
3. School of Biological Sciences University of Adelaide Adelaide South Australia 5005 Australia
4. South Australian Museum Adelaide South Australia 5000 Australia
5. Palaeoscience Research Centre, School of Environmental and Rural Sciences University of New England Armidale New South Wales 2351 Australia