1. Department of Fish, Wildlife, and Conservation Biology Colorado State University Fort Collins Colorado USA
2. Department of Mathematics and Statistics San Diego State University San Diego California USA
3. Denali National Park and Preserve National Park Service Anchorage Alaska USA
4. Arctic Network National Park Service Anchorage Alaska USA
5. Department of Statistics Colorado State University Fort Collins Colorado USA
6. Colorado Cooperative Fish and Wildlife Research Unit U.S. Geological Survey Fort Collins Colorado USA