1. CAS Key Laboratory of Tropical Forest Ecology, Xishuangbanna Tropical Botanical Garden Chinese Academy of Sciences Mengla 666303 Yunnan China
2. Center of Plant Ecology, Core Botanical Gardens, Xishuangbanna Tropical Botanical Garden Chinese Academy of Sciences Mengla 666303 Yunnan China
3. University of Chinese Academy of Sciences Beijing 100049 China
4. Key Laboratory of Vertebrate Evolution and Human Origins, Institute of Vertebrate Paleontology and Paleoanthropology Chinese Academy of Sciences Beijing 100044 China
5. Center for Excellence in Life and Paleoenvironment Chinese Academy of Sciences Beijing 100101 China
6. Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany Chinese Academy of Sciences Kunming 650204 China