1. CAS Key Laboratory of Tropical Forest Ecology Chinese Academy of Sciences Mengla 666303 Yunnan China
2. Center of Plant Ecology, Core Botanical Gardens Chinese Academy of Sciences Mengla 666303 Yunnan China
3. Centre de Recherche en Paléontologie–Paris (CR2P) MNHN ‐ Sorbonne Université–CNRS 43 Rue Buffon Paris France
4. University of Chinese Academy of Sciences Beijing 100049 China
5. CAS Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany Chinese Academy of Sciences Kunming 650204 China
6. School of Environmental, Earth and Ecosystem Sciences The Open University Walton Hall Milton Keynes MK7 6AA UK
7. Key Laboratory of Vertebrate Evolution and Human Origins, Institute of Vertebrate Paleontology and Paleoanthropology Chinese Academy of Sciences Beijing 100044 China