1. School of Forestry and Natural Resources University of Georgia Athens GA 30602 USA
2. Department of Genetics University of Georgia Athens GA 30602 USA
3. College of Forest Resources and Environment Nanjing Forestry University Nanjing 210037 China
4. Institute for Plant Breeding, Genetics and Genomics University of Georgia Athens GA 30602 USA
5. Center for Applied Genetic Technologies University of Georgia Athens GA 30602 USA