1. Department of Integrative Biology University of California Berkeley CA 94720 USA
2. Centre for Ecology and Conservation University of Exeter Penryn Campus Tremough TR11 4EH UK
3. The University of Warwick Coventry CV4 7AL UK
4. Department of Plant Sciences Oxford University Oxford OX1 3RB UK
5. Department of Ecology and Evolutionary Biology Princeton University Princeton NJ 08544 USA