1. State Key Laboratory of Tree Genetics and Breeding, College of Biological Sciences and Technology Beijing Forestry University Beijing 100083 China
2. National Engineering Research Center of Tree Breeding and Ecological Restoration, College of Biological Sciences and Technology Beijing Forestry University No. 35 Qinghua East Road Beijing 100083 China
3. Key Laboratory of Genetics and Breeding in Forest Trees and Ornamental Plants, Ministry of Education, College of Biological Sciences and Technology Beijing Forestry University No. 35, Qinghua East Road Beijing 100083 China
4. The Tree and Ornamental Plant Breeding and Biotechnology Laboratory of National Forestry and Grassland Administration Beijing Forestry University No. 35 Qinghua East Road Beijing 100083 China
5. INRA‐Université de Lorraine, INRAe, UMR 1136, Interactions Arbres/Microorganismes, INRAe‐Grand Est‐Nancy 54280 Champenoux France