1. Systems Ecology Department of Ecological Science Vrije Universiteit Amsterdam De Boelelaan 1085 Amsterdam 1081 HV the Netherlands
2. Grassland Science Department College of Grassland Science and Technology China Agricultural University No. 2 Yuanmingyuan West Road, Haidian District Beijing 100193 China
3. Departamento de Ciencias de la Vida Facultad de Ciencias Universidad de Alcalá Carretera Madrid‐Barcelona Km 33.6 Alcalá de Henares Madrid 28805 Spain
4. Sigur Nature Trust Masinagudi PO Masinagudi Tamil Nadu 643223 India
5. Forest Ecology and Forest Management Group Centre for Ecosystems Wageningen University PO Box 47 Wageningen 6700 AA the Netherlands