1. School of Earth Sciences; Life Sciences Building 24 Tyndall Avenue Bristol BS8 1TQ UK
2. Department of Life Sciences; Natural History Museum; Cromwell Road London SW7 5BD UK
3. Department of Palaeontology; University of Vienna; Geozentrum Althanstrasse 14 Vienna 1090 Austria
4. Chengdu Center of China Geological Survey; Chengdu Institute of Geology & Mineral Resources; Chengdu 610081 China
5. Department of Animal & Plant Sciences; University of Sheffield; Sheffield S10 2TN UK