1. College of Life Sciences Zhejiang University Hangzhou Zhejiang 310058 China
2. Bioinformatics Research Centre Aarhus University C.F. Möllers Allé 8 Aarhus C DK‐8000 Denmark
3. Centre National de la Recherche Scientifique (CNRS) ECOBIO (Ecosystèmes, Biodiversité, Evolution) – Unité Mixte de Recherche (UMR) 6553 Université de Rennes Rennes F‐35000 France
4. Program in Plant Ecology and Evolution Department of Ecology and Genetics Evolutionary Biology Centre Uppsala University Uppsala 75236 Sweden