1. Department of Biological and Environmental Science University of Jyväskylä Jyväskylä Finland
2. Helsinki Institute of Life SciencesUniversity of Helsinki Helsinki Finland
3. Organismal and Evolutionary Biology Research Programme Faculty of Biological and Environmental Sciences University of Helsinki Helsinki Finland
4. Department of Biology Washington University in St. Louis St. Louis MO USA
5. Institute for Biodiversity and Ecosystem Dynamics University of Amsterdam Amsterdam The Netherlands