1. Evolutionary and Behavioural Ecology Department—CEFE, Univ. Montpellier, CNRS, EPHE, IRD , Montpellier , France
2. Organismal and Evolutionary Biology Research Program, Faculty of Biological and Environmental Sciences, University of Helsinki , Helsinki , Finland
3. Department of Biological and Environmental Science, University of Jyväskylä , Jyväskylä , Finland