1. Division of Ecology and Evolution, Research School of BiologyThe Australian National University RN Robertson Building, 46 Sullivans Creek Road Canberra ACT, 2601 Australia
2. Department of Botany, Faculty of ScienceUniversity of Peradeniya Peradeniya 20400 Sri Lanka
3. Medical School, The Australian National University Canberra ACT 2601 Australia
4. Gastroenterology and Hepatology Unit, Canberra Hospital Canberra ACT Australia