1. Ecologie Systématique Evolution, Université Paris‐Sud, CNRS, AgroParisTechUniversité Paris‐Saclay F‐91405 Orsay Cedex France
2. Taps Old Rectory DK‐6040 Christiansfeld Denmark
3. Department of BiologyUniversity of Antwerp Campus Drie Eiken, Building D. D. 137, Universitetsplein 1, B‐2610 Wilrijk Belgium