1. Department of Biology, 510 Mueller Laboratory, Huck Institutes of the Life Sciences The Pennsylvania State University University Park Pennsylvania 16802 USA
2. Key Laboratory of National Forestry and Grassland Administration for Orchid Conservation and Utilization at College of Landscape Architecture Fujian Agriculture and Forestry University Fuzhou 350002 China
3. Shanghai Chenshan Botanical Garden Songjiang Shanghai 201602 China
4. State Key Laboratory of Systematic and Evolutionary Botany, Institute of Botany Chinese Academy of Sciences Beijing 100093 China
5. Fairchild Tropical Botanic Garden Coral Gables Florida 33156 USA