1. College of Horticulture and Landscape Architecture, Zhongkai University of Agriculture and Engineering , Guangzhou 510225 , China
2. Guangxi Nanning Roy Garden Co., Ltd, Nanning 530227, China
3. Shenzhen Key Laboratory of Southern Subtropical Plant Diversity, Fairy Lake Botanical Garden, Shenzhen & Chinese Academy of Sciences , Shenzhen 518004, Guangdong , China
4. State Key Laboratory of Systematic and Evolutionary Botany, Institute of Botany, Chinese Academy of Sciences , Beijing, 100093 , China
5. Key Laboratory of Plant Resources Conservation and Sustainable Utilization, South China Botanical Garden, Chinese Academy of Sciences , Guangzhou 510650 , China
6. College of Forestry and Landscape Architecture, South China Agricultural University , Guangzhou 510642 , China