1. Department of Microbiology, Graduate School of Medical Sciences Kumamoto University Kumamoto Japan
2. Advanced Health Care Science Research Unit, Institute for Frontier Science Initiative Kanazawa University Kanazawa Japan
3. Pulmonary Branch, National Heart, Lung, and Blood Institute National Institutes of Health Bethesda Maryland USA
4. Department of Molecular Infectiology, Graduate School of Medicine Chiba University Chiba Japan