1. Institute of Ocean Sciences, Fisheries and Oceans Canada, Sidney, BC V8L 5T5, Canada
2. Department of Zoology, University of British Columbia, Vancouver, BC V6T 1Z4, Canada
3. Pacific Biological Station, Fisheries and Oceans Canada, Nanaimo, BC V9T 6N7, Canada
4. Department of Mathematics, Simon Fraser University, Burnaby, BC V5A 1S6, Canada
5. Pacific Regional Headquarters, Fisheries and Oceans Canada, Vancouver, BC V6C 3S4, Canada
6. Department of Biology, University of Victoria, Victoria, BC V6T 1Z4, Canada
7. Department of Forest and Conservation Sciences, University of British Columbia, Vancouver, BC V8W 2Y2, Canada