1. US Geological Survey, Northern Rocky Mountain Science Center, 2327 University Way, Suite-2, Bozeman, MT 59715, USA.
2. US Geological Survey, Forest and Rangeland Ecosystem Science Center, 970 Lusk Street, Boise 83706, USA.
3. Montana Cooperative Fishery Research Unit, Department of Ecology, Montana State University, P.O. Box 173460, Bozeman, MT 59717, USA.
4. Montana Fish, Wildlife, and Parks, 3201 Spurgin Road, Missoula, MT 59804, USA.
5. Confederated Tribes of the Colville Reservation, Fish and Wildlife Department, Okanogan Basin Monitoring and Evaluation Program, Nespelem, WA 99155, USA.
6. US Geological Survey, Montana Cooperative Fishery Research Unit, Department of Ecology, Montana State University, PO Box 173460, Bozeman, MT 59717, USA.