1. Tour du Valat, 55266, Research Institute for the Conservation of Mediterranean Wetlands, Species department, Arles, Provence-Alpes-Côte d'Azu, France
2. UMR ECOBIOP, UMR 1224, inrae, university of Pau & Pays de l'Adour, 64310, Saint-Pée sur Nivelle, France
3. University of California Berkeley, 1438, Department of Environmental Science, Policy & Management, Berkeley, California, United States;
4. UMR ECOBIOP, UMR 1224, inrae, university of Pau & Pays de l'Adour, 64310, Saint-Pée sur Nivelle, France;
5. Université de Pau et des Pays de l'Adour, 27073, EcoBiop, 173 RD 918, Route de Saint-Jean de Luz, Saint-Pée sur Nivelle, France, 64310;
6. University of California - Berkeley, Department of Environmental Science, Policy & Management, Berkeley, California, United States;