1. a Centre for Ecology and Hydrology, Wallingford, Oxfordshire OX10 8BB, UK
2. b Heath Barton, Manor Road, Goring-on-Thames, Oxfordshire RG8 9EH, UK
3. c School of Development Studies, University of East Anglia, Norwich NR4 7TJ, UK
4. d Centre for Ecology and Hydrology, Wallingford, Oxfordshire OX10 8BB, UK