1. Iwate Biotechnology Research Center, 22-174-4, Narita, Kitakami, Iwate 024-0003, Japan
2. Department of Biotechnology and Life Science, Faculty of Engineering, Tokyo University of Agriculture and Technology, 2-24-16, Koganei, Tokyo 184-8588, Japan
3. Faculty of Agriculture, Iwate University, 3-18-8, Ueda, Morioka 020-8550, Japan
4. Department of Biological and Environmental Science, Graduate School of Agriculture, Shizuoka University, 836, Ohya, Suruga, Shizuoka 422-8529, Japan