1. School of Life Sciences, University of Warwick, Coventry CV4 7AL, UK
2. Present address: Environment and Sustainability Institute, University of Exeter, Penryn Campus, Penryn, Cornwall, TR10 9FE, UK.
3. Present address: The Native Antigen Company, Langford Locks, Kidlington, Oxford, OX5 1LH, UK.
4. Present address: Waters Corporation, Stamford Avenue, Altrincham Road, Wilmslow, SK9 4AX, UK.