1. School of Earth and Environmental Sciences and Research Institute of Oceanography, Seoul National University, Seoul 08826, Republic of Korea
2. Division of Life Sciences, Korea Polar Research Institute, 26 Songdomirae-ro, Yeonsu-gu, Incheon 21990, Republic of Korea
3. Division of Microbiology, Honam National Institute of Biological Resources, 99 Gohadoan-gil, Mokpo-si, Jeollanam-do, Republic of Korea
4. Division of Earth-System Sciences, Korea Polar Research Institute, 26 Songdomirae-ro, Yeonsu-gu, Incheon 21990, Republic of Korea