1. Plant and Invertebrate Ecology Department, Rothamsted Research, Harpenden, Hertfordshire AL5 2JQ, United Kingdom;
2. School of Physical, Environmental and Mathematical Sciences, The University of New South Wales at the Australian Defence Force Academy, Canberra, ACT 2600, Australia;
3. Institute of Applied Ecology, University of Canberra, ACT 2601, Australia
4. Natural Resources Institute, University of Greenwich, Chatham, Kent ME4 4TB, United Kingdom;