1. Institut Méditerranéen de Biodiversité et d’Ecologie, Avignon Université, CNRS, IRD, Aix Marseille Université, 84911 Avignon, France.
2. Centre for African Ecology, School of Animal, Plant and Environmental Sciences, University of the Witwatersrand, Johannesburg 2050, South Africa.
3. Instituto de Biociências, Lab of Vegetation Ecology, Universidade Estadual Paulista (UNESP), Rio Claro 13506-900, Brazil.
4. Department of Ecology and Evolutionary Biology, University of Colorado, Boulder, CO, USA.
5. Institute of Arctic and Alpine Research, University of Colorado, Boulder, CO, USA.